Below is a list of my peer-reviewed articles and preprints, with links to DOIs of all open access materials and to files of the few closed access works. These files and a "standalone" list showing the DOIs and links in full (also archived with DOI 10.5281/zenodo.4730019) are at

If you have any trouble accessing anything, or need further data or information, do not hesitate to contact me.

Peer-reviewed articles

Sookias RB, Dilkes D, Sobral G, Smith RMH, Wolvaardt FP, Arcucci AB, Bhullar B-AS, Werneburg I. 2020. The craniomandibular anatomy of the early archosauriform Euparkeria capensis and the dawn of the archosaur skull. Royal Society Open Science. DOI (publisher open access).

Sookias RB. 2019. Exploring the effects of character construction and choice, outgroups, and analytical method on phylogenetic inference from discrete characters in extant crocodilians. Zoological Journal of the Linnean Society. DOI (publisher closed access), main text (pdf), supplement (zip). I have also made an additional spreadsheet highlighting differences in scoring of Bernissartia from the matrix of Narváez et al. 2016, and provide all my photographs of Bernissartia here (zip).

Butler RJ, Ezcurra MD, Liu J, Sookias RB, Sullivan C. 2019. The anatomy and phylogenetic position of the erythrosuchid archosauriform Guchengosuchus shiguaiensis from the earliest Middle Triassic of China. PeerJ 7:e6435 . DOI (publisher open access).

Sookias RB, Passmore S, Atkinson QD. 2018. Deep cultural ancestry and human development indicators across nation states. Royal Society Open Science. DOI (publisher open access).

Foth C, Ezcurra MD, Sookias RB, Brusatte SL, Butler RJ. 2016. Unappreciated diversification of stem archosaurs during the Middle Triassic predated the dominance of dinosaurs. BMC Evolutionary Biology 16:188. DOI (publisher open access).

Sobral G, Sookias RB, Bhullar B-AS, Smith R, Butler RJ, Müller J. 2016. New information on the braincase and inner ear of Euparkeria capensis Broom: implications for diapsid and archosaur evolution. Royal Society Open Science 3:160072. DOI (open access). N.B. Supplementary figures do not appear to be downloading correctly - I have thus made them available here.

Sookias RB. 2016. The relationships of the Euparkeriidae and the rise of Archosauria. Royal Society Open Science 3:150674. DOI (publisher open access).

Sookias RB, Sullivan, C., Liu, J. & Butler, R. J. 2014. Systematics of putative euparkeriids (Diapsida: Archosauriformes) from the Triassic of China. PeerJ. 2:e658. DOI (publisher open access).

Sookias RB, Böhmer C, Clack JA. 2014. Redescription and phylogenetic analysis of an enigmatic Pennsylvanian (Late Carboniferous) tetrapod from Nova Scotia, and the lability of Meckelian jaw ossification. PLoS ONE 9:e109717. DOI (publisher open access).

Sookias RB, Sennikov AG, Gower, DJ, Butler RJ. 2014. The monophyly of Euparkeriidae (Reptilia: Archosauriformes) and the origins of crown Archosauria: a revision of Dorosuchus neoetus from the Middle Triassic of Russia. Palaeontology 57: 1177-1202. DOI (publisher open access).

Butler RJ, Sullivan C, Ezcurra MD, Liu J, Lecuona A, Sookias RB. 2014. New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation. BMC Evolutionary Biology 14:128. DOI (publisher open access).

Sookias RB, Budd GE, Kear BP. 2013. Mesozoic fossil sustainability: synoptic case studies of resource management. GFF 135:131-143. DOI (publisher closed access), main text (pdf), supplement (ods).

Sookias RB, Butler RJ. 2013. Euparkeriidae. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379:35-48. DOI (publisher closed access), main text (pdf).

Sookias RB, Benson RBJ, Butler RJ. 2012. Biology, not environment, drives major patterns in maximum tetrapod body size through time. Biology Letters 8:674-677. DOI (publisher open access)

Sookias RB, Butler RJ, Benson RBJ. 2012 Rise of dinosaurs reveals major body size transitions are driven by passive processes of trait evolution. Proceedings of the Royal Society B 279:2180-2187. DOI (publisher open access).

Non-peer-reviewed preprints

Sookias RB. 2019, Feb. 13. The concept of homoiology and its potential application in palaeontological phylogenetics. PaleorXiv. 

© Images copyright 2019 Cam-Anh Luong

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